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Aβ40 and Aβ42 make separate fibrils after mixed nuclei

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In mixtures, Aβ42 fibrillizes first; elongation and secondary nucleation stay homospecific, so each peptide builds its own fibrils.

Source

The Aβ40 and Aβ42 peptides self-assemble into separate homomolecular fibrils in binary mixtures but cross-react during primary nucleation

Cukalevski R, Yang X, Meisl G, et al. · Chemical science · 2015

doi.org/10.1039/c4sc02517bRead the full paper ↗140 citationscc by

What they did

Authors combined ThT, CD, isotope-resolved MS/NMR, cryo-TEM pitch measurements and self- vs cross-seeding to dissect microscopic steps in Aβ40:Aβ42 mixtures.

What they found

Two transitions below ~10 μM; first fibrils are Aβ42 (<13% Aβ40). Helical half-pitch 162 nm (Aβ40) vs 31 nm (Aβ42). Cross-seeding is weak versus self-seeding; Aβ42 monomers accelerate Aβ40 primary nucleation.

The limits

What it doesn't show

Atomic mixed-oligomer structures are not solved; disease relevance is inferred from peptide biophysics, not patients.

Key terms

Primary nucleation
Formation of the first aggregates from monomers; the least sequence-specific step here.
Secondary nucleation
Surface-catalyzed formation of new fibrils on existing fibrils; highly homospecific.
Homomolecular fibril
Fibril built from only Aβ40 or only Aβ42, not a statistical copolymer.
Thioflavin T (ThT)
Dye whose fluorescence reports β-sheet amyloid.
Cross-seeding
Using fibrils of one peptide to nucleate the other.

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Quiz yourself

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Fibrils in Aβ40/Aβ42 mixtures are:

Common questions

Do mixed fibrils form?

Not detectably; fibrils are homomolecular.

Where do the peptides mix?

At primary nucleation/small oligomers.

Which peptide aggregates first?

Aβ42, then Aβ40.

How do TEM pitches differ?

~162 nm (Aβ40) vs ~31 nm (Aβ42).

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