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Does vagal TLR4 cause LPS anorexia—or mainly CGRP release?

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Restoring TLR4 only in Nav1.8 afferents did not restore LPS anorexia, but LPS still triggered CGRP release from TLR4+/CGRP+ vagal jugular neurons.

Source

TLR4 Signaling Selectively and Directly Promotes CGRP Release from Vagal Afferents in the Mouse

Jia L, Lee S, Tierney JA, et al. · eNeuro · 2021

doi.org/10.1523/eneuro.0254-20.2020Read the full paper ↗41 citationscc by

What they did

Built Tlr4 LoxTB mice, restored TLR4 globally (Zp3-Cre) or in Nav1.8 afferents, mapped Tlr4 with ISH, and applied LPS to cultured afferents.

What they found

Nav1.8-restricted TLR4 was insufficient for rapid-onset LPS anorexia; Tlr4 enriched in jugular Nav1.8/CGRP vagal afferents; LPS stimulated CGRP release in vitro.

The limits

What it doesn't show

Does not map the full in vivo infection-fighting role of the LPS–TLR4–CGRP loop or test female experimental cohorts for feeding assays.

Key terms

TLR4
Innate immune receptor that senses bacterial LPS.
LPS
Lipopolysaccharide from Gram-negative bacteria.
CGRP
Calcitonin gene-related peptide released from sensory afferents.
Nav1.8 afferents
Peripheral sensory neurons expressing the Nav1.8 sodium channel.
Tlr4 LoxTB
Mouse with a loxP-flanked transcription blocker silencing Tlr4 until Cre removal.
Jugular ganglion
Vagal sensory ganglion enriched for Tlr4/CGRP neurons here.

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Quiz yourself

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Nav1.8-restricted TLR4 restored LPS anorexia?

Common questions

Did Nav1.8-TLR4 restore anorexia?

No—LPS did not cause rapid-onset anorexia.

What did LPS trigger?

CGRP release from cultured Nav1.8-restricted TLR4 afferents.

Where is Tlr4 enriched?

Jugular vagal Nav1.8 afferents co-expressing CGRP.

New tool?

Cre-reactivatable Tlr4 LoxTB mouse.