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How can flies adapt via many interchangeable alleles?

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Ten D. simulans replicates converged phenotypically under a new temperature regime, but used largely different selected alleles—evidence for genetic redundancy.

Source

Genetic redundancy fuels polygenic adaptation in Drosophila

Barghi N, Tobler R, Nolte V, et al. · PLoS biology · 2019

doi.org/10.1371/journal.pbio.3000128Read the full paper ↗272 citationscc by

What they did

Used laboratory natural selection on 10 replicate Drosophila simulans populations under a new temperature regime, tracking phenotypes and allele-frequency changes for selective sweeps vs polygenic adaptation.

What they found

Convergent fitness/metabolic/fat phenotypes with a strong polygenic response (99 selected alleles; mean s = 0.059), yet each allele rose only in a subset of replicates. Redundancy/QT paradigms fit better than independent selective-sweep simulations.

The limits

What it doesn't show

Lab temperature selection is not a direct map of wild climate adaptation; molecular non-convergence does not mean phenotypes lack shared physiological mechanisms.

Key terms

Polygenic adaptation
Adaptation via frequency shifts at many loci rather than one large-effect sweep.
Genetic redundancy
Different beneficial alleles can substitute for one another across replicates.
AFC
Allele frequency change used to detect selected variants.
Selective sweep
Rapid rise of a strongly favored allele, often leaving a narrow genomic signature.
Quantitative trait paradigm
Many small-effect alleles push a trait toward a shared optimum.
Laboratory natural selection
Replicated experimental evolution under controlled selective conditions.

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Number of evolving replicates:

Common questions

How many replicates?

10 evolving D. simulans populations.

How many selected alleles flagged?

99 selected alleles (mean s = 0.059).

Did genetics converge across replicates?

No—alleles rose only in subsets of replicates despite phenotype convergence.

What paradigms fit best?

Redundancy and quantitative-trait paradigms beat independent sweep models.

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