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Biodiversity · Taxonomy

About 8.7 million species — and roughly seven in eight have never been described

Evidence: EmergingMore than one study points the same way, but the body is still thin. What the labels mean

Study published Jan 1, 2011. PaperFren added this explanation Sep 20, 2026.

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Short answer

A higher-taxon extrapolation estimates ~8.7 million eukaryotic species, of which roughly 14% on land and 9% in the ocean are described.

What happened

Mora, Tittensor, Adl, Simpson and Worm used the fact that higher taxa (phyla, classes, orders) are discovered and then saturate long before species do. Fitting that pattern and extrapolating gives ~8.7 million eukaryotic species on Earth and ~2.2 million in the ocean. Validation against well-sampled groups supported the approach. A parallel survey drew responses from 548 of about 2,938 contacted taxonomists on synonymy and effort.

Why it matters

The headline is not the number but the fraction: on this estimate only about 14% of terrestrial and 9% of marine eukaryotic species have been catalogued after 250 years of taxonomy. That sets the scale of what biodiversity loss statistics are actually measuring.

Evidence

Study type
Higher-taxon extrapolation model validated against well-sampled groups, with a taxonomist survey
Sample
Global taxonomic catalogues; 548 of ~2,938 contacted taxonomists responded (18.7%)
Journal
PLoS Biology · peer reviewed
Replication
The approach has been re-run and contested by later estimates; the order of magnitude is more durable than the exact figure
Limitations
A model, not a census. Prokaryotes are excluded because the species concept does not transfer. Unresolved synonyms bias the catalogued baseline.

What this connects to

Sources

The 2 studies this explanation is built from, by the role each plays. Every source links to PaperFren’s explanation of it and to the original paper.

Primary study

Supporting evidence

Before

Published expert estimates of global species richness spanned more than an order of magnitude, from about 3 million to 100 million, with no shared method to choose between them.

Now

A method with an internal validity check narrows the range considerably. It is still a model extrapolation, it does not cover prokaryotes — whose species concept differs — and it will move as synonymy and sampling improve.